But Firestar's dreams are haunted by wailing cats fleeing a terrible disaster. With unexpected help from an old kittypet friend, he discovers a shocking secret: StarClan, the warrior ancestors who guide his paw steps, have lied to him.

Firestar is faced with the hardest decision of his life. Can he really turn his back on the forest that has become his home and embark on a perilous quest to discover a dark truth—one that has been buried beyond the memory of living cats? Whatever he finds at the end of his journey, he knows that nothing can ever be the same again.