Centuries have passed since the forces of nature won the war against humanity. Sentient animals now rule a healing world, and as the stain of mankind continues to dwindle, a young wolf called Silversong is determined to rise in the hierarchy of his pack. Strong at manipulating wind and air, all he needs is a way to prove himself to his Chief.

Before he can get the respect he deserves, however, Silversong's aspirations are cut short by the Heretic and his outcast wolves. Against all odds, the Heretic and his band of exiles escape their imprisonment far to the west and wreak havoc on Silversong's pack. The exiles pose a threat unlike any other, and their enigmatic leader won't stop his brutal conquest until all wolfkind submits to him.

Silversong can't let a monstrous wolf like the Heretic roam free. With the wind at his back, he pursues the leader of the exiles into forests of shadow and into ancient places better left forgotten. But the further he strays from home, the more he comes to realize that maybe his enemies aren't so evil after all. Maybe there's a reason for the destruction they seek... and maybe there's a far greater danger lying in wait.